Hymenasplenium
| Hymenasplenium | |
|---|---|
 | |
| Hymenasplenium cheilosorum | |
Scientific classification  | |
| Kingdom: | Plantae |
| Clade: | Embryophytes |
| Clade: | Tracheophytes |
| Division: | Polypodiophyta |
| Class: | Polypodiopsida |
| Order: | Polypodiales |
| Suborder: | Aspleniineae |
| Family: | Aspleniaceae |
| Genus: | Hymenasplenium Hayata |
| Type species | |
| Hymenasplenium unilaterale (Lam.) Hayata. 1927 | |
| Synonyms | |
| |
Hymenasplenium is one of three genera of ferns in the Aspleniaceae (spleenwort family), in the eupolypods II clade of the order Polypodiales.[1][2] The others are Hemidictyum and Asplenium. Hymenasplenium was segregated because it is a natural grouping with differing rhizome morphology – dorsiventral v. radial for the rest of Asplenium, differing chromosome count – x=39 v. x=36 for the rest of Asplenium, and a clear monophyletic grouping based on genetic analysis. It was confirmed as a sister group to Asplenium in a 2015 molecular study of the genera.[3]
List of species
[edit]As of July 2025[update], the Checklist of Ferns and Lycophytes of the World recognized the sixty-seven species and three hybrids below.[4]
- Hymenasplenium amoenum (C.Presl ex Mett.) Ranil
- Hymenasplenium andohahelense Viane
- Hymenasplenium antsirananense Viane
- Hymenasplenium apogamum (N.Murak. & Hatan.) Nakaike
- Hymenasplenium basiscopicum (R.C.Moran & Sundue) L.Regalado & Prada
- Hymenasplenium cardiophyllum (Hance) Nakaike
- Hymenasplenium changputungense (Ching) Viane & S.Y.Dong
- Hymenasplenium cheilosorum (Kunze ex Mett.) Tagawa
- Hymenasplenium chingii K.W.Xu, Li Bing Zhang & W.B.Liao
- Hymenasplenium delitescens (Maxon) L.Regalado & Prada
- Hymenasplenium denticulatum K.W.Xu, Li Bing Zhang & W.B.Liao
- Hymenasplenium distans Li Bing Zhang, K.W.Xu & Liang Zhang
- Hymenasplenium excisum (C.Presl) S.Linds.
- Hymenasplenium filipes (Copel.) Sugim.
- Hymenasplenium furfuraceum (Ching) Viane & S.Y.Dong
- Hymenasplenium guineense Li Bing Zhang, W.B.Liao & K.W.Xu
- Hymenasplenium hastifolium K.W.Xu, Li Bing Zhang & W.B.Liao
- Hymenasplenium hoffmannii (Hieron.) L.Regalado & Prada
- Hymenasplenium hondoense (N.Murak. & Hatan.) Nakaike
- Hymenasplenium hovenkampii K.W.Xu & Li Bing Zhang
- Hymenasplenium ikenoi (Makino) Viane
- Hymenasplenium inthanonense N.Murak. & J.Yokoy.
- Hymenasplenium kenyense Li Bing Zhang, K.W.Xu & Kamau
- Hymenasplenium kinabaluense Li Bing Zhang & K.W.Xu
- Hymenasplenium laetum (Sw.) L.Regalado & Prada
- Hymenasplenium latedens (Ching) Viane & S.Y.Dong
- Hymenasplenium laterepens N.Murak. & X.Cheng ex Y.Fen Chang & K.Hori
- Hymenasplenium madagascariense Li Bing Zhang, W.B.Liao & K.W.Xu
- Hymenasplenium murakami-hatanakae Nakaike
- Hymenasplenium neocaledonicum Li Bing Zhang & K.W.Xu
- Hymenasplenium ngheanense Li Bing Zhang, K.W.Xu & N.T.Lu
- Hymenasplenium nigricostatum Li Bing Zhang & K.W.Xu
- Hymenasplenium obliquissimum (Hayata) Sugim.
- Hymenasplenium obscurum (Blume) Tagawa
- Hymenasplenium obtusidentatum Y.Fen Chang & G.Cheng Zhang
- Hymenasplenium obtusifolium (L.) L.Regalado & Prada
- Hymenasplenium ofeliae (Salgado) Barcelona & Pelser
- Hymenasplenium oligosorum Li Bing Zhang, K.W.Xu & Lorence
- Hymenasplenium ortegae (N.Murak. & R.C.Moran) L.Regalado & Prada
- Hymenasplenium perriei Li Bing Zhang & K.W.Xu
- Hymenasplenium phamhoanghoi Li Bing Zhang, K.W.Xu & T.T.Luong
- Hymenasplenium praestans (Copel.) Testo, Riibe & Sundue
- Hymenasplenium pseudobscurum Viane
- Hymenasplenium pubirhizoma (Ching & Z.Y.Liu) K.W.Xu & Li Bing Zhang
- Hymenasplenium purpurascens (Mett.) L.Regalado & Prada
- Hymenasplenium quangnamense Li Bing Zhang, K.W.Xu & Liang Zhang
- Hymenasplenium queenslandicum Li Bing Zhang & K.W.Xu
- Hymenasplenium quercicola (Ching) Viane & S.Y.Dong
- Hymenasplenium repandulum (Kunze) L.Regalado & Prada
- Hymenasplenium riparium (Liebm.) L.Regalado & Prada
- Hymenasplenium rivulare (Fraser-Jenk.) Viane & S.Y.Dong
- Hymenasplenium sabahense Li Bing Zhang, K.W.Xu & C.W.Chen
- Hymenasplenium samoaense Li Bing Zhang, K.W.Xu & K.R.Wood
- Hymenasplenium sinense K.W.Xu, Li Bing Zhang & W.B.Liao
- Hymenasplenium solomonense Li Bing Zhang & K.W.Xu
- Hymenasplenium speluncicola Li Bing Zhang, K.W.Xu & H.He
- Hymenasplenium subnormale (Copel.) Nakaike
- Hymenasplenium szechuanense (Ching) Viane & S.Y.Dong
- Hymenasplenium tholiforme Liang Zhang, W.B.Ju & K.W.Xu
- Hymenasplenium trapeziforme (Wall. ex Roxb.) comb.ined., Asplenium trapeziforme Wall. ex Roxb.
- Hymenasplenium triquetrum (N.Murak. & R.C.Moran) L.Regalado & Prada
- Hymenasplenium unilaterale (Lam.) Hayata
- Hymenasplenium vanuatuense Li Bing Zhang & K.W.Xu
- Hymenasplenium volubile (N.Murak. & R.C.Moran) L.Regalado & Prada
- Hymenasplenium wangpeishanii Li Bing Zhang & K.W.Xu
- Hymenasplenium wildii (Bail.) D.J.Ohlsen
- Hymenasplenium wusugongii Li Bing Zhang, W.B.Liao & K.W.Xu
Hybrids
[edit]- Hymenasplenium × doanyense Viane
- Hymenasplenium × excurum Viane
- Hymenasplenium × incisoserratum (Rosenst.) comb.ined., Asplenium × incisoserratum (Rosenst.) N.Murak. & R.C.Moran
Phylogeny
[edit]| Phylogeny from Fern Tree of Life[5][6] | |||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
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References
[edit]- ↑ Alan R. Smith; Kathleen M. Pryer; Eric Schuettpelz; Petra Korall; Harald Schneider; Paul G. Wolf (2006). "A classification for extant ferns" (PDF). Taxon. 55 (3): 705–731. Bibcode:2006Taxon..55..705S. doi:10.2307/25065646. JSTOR 25065646. Archived from the original (PDF) on February 26, 2008.
- ↑ Maarten J. M. Christenhusz; Xian-Chun Zhang; Harald Schneider (2011). "A linear sequence of extant families and genera of lycophytes and ferns" (PDF). Phytotaxa. 19: 7–54. Bibcode:2011Phytx..19....7C. doi:10.11646/phytotaxa.19.1.2.
- ↑ Ohlsen DJ, Perrie LR, Shepherd LD, Brownsey PJ, Bayly MJ (2015). "Phylogeny of the fern family Aspleniaceae in Australasia and the south-western Pacific". Australian Systematic Botany. 27 (6): 355–71. doi:10.1071/sb14043.
- ↑ Hassler, Michael. "Hymenasplenium". World Ferns. Retrieved 2025-07-15.
- ↑ Nitta, Joel H.; Schuettpelz, Eric; Ramírez-Barahona, Santiago; Iwasaki, Wataru; et al. (2022). "An Open and Continuously Updated Fern Tree of Life". Frontiers in Plant Science. 13 909768. Bibcode:2022FrPS...1309768N. doi:10.3389/fpls.2022.909768. PMC 9449725. PMID 36092417.
- ↑ "Tree viewer: interactive visualization of FTOL". FTOL v1.7.0 (GenBank release 261). 2024. Retrieved 1 May 2025.
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