Heteronychus

Heteronychus
Heteronychus arator
Scientific classification
Kingdom:	Animalia
Phylum:	Arthropoda
Clade:	Pancrustacea
Class:	Insecta
Order:	Coleoptera
Suborder:	Polyphaga
Infraorder:	Scarabaeiformia
Family:	Scarabaeidae
Tribe:	Pentodontini
Genus:	Heteronychus Dejean, 1833

Heteronychus is a genus of beetles belonging to the family Scarabaeidae, subfamily Dynastinae.

The species of this genus are found in Africa, Southeastern Asia and Australia.

Species:

• Heteronychus abyssinicus Jack, 1923
• Heteronychus amplipennis Benderitter, 1915
• Heteronychus amplus Kolbe, 1900
• Heteronychus andersoni Jack, 1923
• Heteronychus annulatus Bates, 1891
• Heteronychus approximans Kolbe, 1900
• Heteronychus arator (Fabricius, 1775)
• Heteronychus ascanius Kolbe, 1900
• Heteronychus atratus Klug, 1855
• Heteronychus basilewskyi Endrödi, 1962
• Heteronychus bituberculatus Kolbe, 1900
• Heteronychus brittoni Ferreira, 1965
• Heteronychus carvalhoi Ferreira, 1970
• Heteronychus citernii Paoli, 1934
• Heteronychus congoensis Kolbe, 1900
• Heteronychus consimilis Kolbe, 1900
• Heteronychus cordatus Endrödi, 1974
• Heteronychus costatus Lansberge, 1886
• Heteronychus cricetus (Hausmann, 1807)
• Heteronychus curtulus Fairmaire, 1887
• Heteronychus desaegeri Burgeon, 1947
• Heteronychus digitiformis Zhang, 1983
• Heteronychus fossor Reiche, 1847
• Heteronychus howdeni Yamaya, 2016
• Heteronychus impudens Jack, 1923
• Heteronychus infans Kolbe, 1897
• Heteronychus inoportunus Ferreira, 1966
• Heteronychus insignificus Jack, 1923
• Heteronychus intermedius Zhang, 1983
• Heteronychus jacki Arrow, 1936
• Heteronychus krombeini Endrödi, 1976
• Heteronychus licas (Klug, 1835)
• Heteronychus lioderes Redtenbacher, 1867
• Heteronychus lusingae Endrödi, 1974
• Heteronychus minimus Dupuis, 2012
• Heteronychus minutus Burmeister, 1847
• Heteronychus mollis Endrödi, 1961
• Heteronychus monodi Paulian, 1954
• Heteronychus mosambicus Péringuey, 1908
• Heteronychus muticus Benderitter, 1915
• Heteronychus paolii Arrow, 1936
• Heteronychus parumpunctatus Burmeister, 1847
• Heteronychus parvus Burmeister, 1847
• Heteronychus pauperatus Péringuey, 1901
• Heteronychus plebejus (Klug, 1832)
• Heteronychus puerilis Kolbe, 1900
• Heteronychus puncticollis Jack, 1923
• Heteronychus punctolineatus Fairmaire, 1893
• Heteronychus pygidialis Kolbe, 1900
• Heteronychus rusticus (Klug, 1832)
• Heteronychus sabackyi Endrödi, 1975
• Heteronychus sacchari Arrow, 1908
• Heteronychus similis Dechambre, 1992
• Heteronychus simulans Jack, 1923
• Heteronychus sublaevis (Fairmaire, 1891)
• Heteronychus tenuestriatus Fairmaire, 1893
• Heteronychus tesari Endrödi, 1968
• Heteronychus tristis Boheman, 1857
• Heteronychus vixstriatus Jack, 1923
• Heteronychus wittei Janssens, 1942