Geomitridae

Geomitridae
	Cernuella virgata
Scientific classification
Kingdom:	Animalia
Phylum:	Mollusca
Class:	Gastropoda
Order:	Stylommatophora
Superfamily:	Helicoidea
Family:	Geomitridae; Boettger, 1909
Type genus
	Geomitra; Swainson, 1840
Genera
	See text

Geomitridae is a taxonomic family of small to medium-sized air-breathing land snails, terrestrial pulmonate gastropod mollusks in the superfamily Helicoidea.^[1]^[2]

Anatomy

The family is characterized by the presence of a free right ommatophoral retractor (passing outside the peni-oviducal angle) and a double stimulatory apparatus.^[1] A free right ommatophoral retractor has been linked to adaptation to xeric habitats.^[3]

Taxonomy

The family Geomitridae is subdivided in the following subfamilies (according to Razkin et al., 2015^[1]). Genera with the family Geomitridae include:^[1]^[4]

Family Geomitridae

Not belonging to a subfamily:
- Keraea Gude, 1911
- Peridotitea Torres Alba, Holyoak, D. T., Holyoak, G. A., Vázquez Toro & Ripoll, 2018

Subfamily Geomitrinae
- tribe Cochlicellini Schileyko, 1972
  - Cochlicella Férussac, 1821
  - Monilearia Mousson, 1872
  - Obelus W. Hartmann, 1842
  - Ripkeniella Hutterer & E. Gittenberger, 1998
- tribe Geomitrini C. Boettger, 1909 - synonym: Ochthephilinae Zilch, 1960 (n.a.)
  - Actinella R. T. Lowe, 1852^[5]
  - Callina R. T. Lowe, 1855
  - Caseolus R. T. Lowe, 1852^[5]
  - Discula R. T. Lowe, 1852^[5]
  - Disculella Pilsbry, 1895^[5]
  - Domunculifex Brozzo, De Mattia, Harl & Neiber, 2020
  - Geomitra Swainson, 1840^[5]
  - Helicomela R. T. Lowe, 1855
  - Heterostoma W. Hartmann, 1843:^[5] synonym of Steenbergia Mandahl-Barth, 1950 (invalid: junior homonym of Heterostoma de Filippi, 1837 [Platyhelminthes])
  - Hystricella R. T. Lowe, 1855^[5]
  - Lemniscia R. T. Lowe, 1855^[5]
  - Moreletina de Frias Martins, 2002^[5]
  - Plebecula R. T. Lowe, 1852
  - Pseudocampylaea L. Pfeiffer, 1877^[5]
  - Serratorotula Groh & Hemmen, 1986^[5]
  - Spirorbula R. T. Lowe, 1852^[5]
  - Steenbergia Mandahl-Barth, 1950
  - Testudodiscula Brozzo, De Mattia, Harl & Neiber, 2020
  - Wollastonaria De Mattia, Neiber & Groh, 2018
- tribe Ponentinini Razkin et al., 2015
  - Ponentina P. Hesse, 1921

subfamily Helicellinae
- tribe Cernuellini Schileyko, 1991
  - Alteniella Clerx & Gittenberger, 1977
  - Cernuella Schlüter, 1838
  - Microxeromagna Ortiz de Zárate López, 1950
  - Xeroplana Monterosato, 1892
  - Xerosecta Monterosato, 1892
- tribe Helicellini Ihering, 1909 - synonym: Jacostidae Pilsbry, 1948 (inv.)
  - Backeljaia Chueca et al., 2018
  - Candidula Kobelt, 1871
  - Helicella Férussac, 1821
  - Orexana Chueca et al., 2018
  - Xerogyra Monterosato, 1892
  - Xeroleuca Kobelt, 1877
  - Xeroplexa Monterosato, 1892
  - Xerotricha Monterosato, 1892
  - Zarateana Chueca et al., 2018
- tribe Helicopsini H. Nordsieck, 1987
  - Helicopsis Fitzinger, 1833
  - Pseudoxerophila Westerlund, 1879
  - Xerolenta Monterosato, 1892
  - Xeromunda Monterosato, 1892
  - Xeropicta Monterosato, 1892
- tribe Plentuisini Razkin et al., 2015
  - Plentuisa Puente & Prieto, 1992
- tribe Trochoideini H. Nordsieck, 1987
  - Trochoidea T. Brown, 1827
  - Xerocrassa Monterosato, 1892

References

1 2 3 4 Razkin, Oihana; Gómez-Moliner, Benjamín Juán; Prieto, Carlos Enrique; Martínez-Ortí, Alberto; Arrébola, José Ramón; Muñoz, Benito; Chueca, Luis Javier; Madeira, María José (February 2015). "Molecular phylogeny of the western Palaearctic Helicoidea (Gastropoda, Stylommatophora)". Molecular Phylogenetics and Evolution. 83: 99–117. doi:10.1016/j.ympev.2014.11.014. PMID 25485783.
↑ "WoRMS - World Register of Marine Species - Geomitridae C. R. Boettger, 1909". www.marinespecies.org. Retrieved 5 September 2024.
↑ Giusti, F; Manganelli, G (1987). "Notulae malacologicae, XXXVI. On some Hygromiidae (Gastropoda: Helicoidea) living in Sardinia and in Corsica.(Studies on the Sardinian and Corsican malacofauna VI)". Bollettino Malacologico. 23: 123–206.
↑ Chueca, Luis J.; Gómez-Moliner, Benjamín J.; Madeira, María José; Pfenninger, Markus (January 2018). "Molecular phylogeny of Candidula (Geomitridae) land snails inferred from mitochondrial and nuclear markers reveals the polyphyly of the genus". Molecular Phylogenetics and Evolution. 118: 357–368. doi:10.1016/j.ympev.2017.10.022. ISSN 1055-7903. PMID 29107619.
1 2 3 4 5 6 7 8 9 10 11 12 "Geomitrini". Fauna Europaea, last update 27 January 2011, accessed 4 August 2011.

Bouchet P., Rocroi J.P., Hausdorf B., Kaim A., Kano Y., Nützel A., Parkhaev P., Schrödl M. & Strong E.E. (2017). Revised classification, nomenclator and typification of gastropod and monoplacophoran families. Malacologia. 61(1-2): 1-526

[:0-1] 1 2 3 4 Razkin, Oihana; Gómez-Moliner, Benjamín Juán; Prieto, Carlos Enrique; Martínez-Ortí, Alberto; Arrébola, José Ramón; Muñoz, Benito; Chueca, Luis Javier; Madeira, María José (February 2015). "Molecular phylogeny of the western Palaearctic Helicoidea (Gastropoda, Stylommatophora)". Molecular Phylogenetics and Evolution. 83: 99–117. doi:10.1016/j.ympev.2014.11.014. PMID 25485783.

[WoRMSD-2] "WoRMS - World Register of Marine Species - Geomitridae C. R. Boettger, 1909". www.marinespecies.org. Retrieved 5 September 2024.

[3] Giusti, F; Manganelli, G (1987). "Notulae malacologicae, XXXVI. On some Hygromiidae (Gastropoda: Helicoidea) living in Sardinia and in Corsica.(Studies on the Sardinian and Corsican malacofauna VI)". Bollettino Malacologico. 23: 123–206.

[4] Chueca, Luis J.; Gómez-Moliner, Benjamín J.; Madeira, María José; Pfenninger, Markus (January 2018). "Molecular phylogeny of Candidula (Geomitridae) land snails inferred from mitochondrial and nuclear markers reveals the polyphyly of the genus". Molecular Phylogenetics and Evolution. 118: 357–368. doi:10.1016/j.ympev.2017.10.022. ISSN 1055-7903. PMID 29107619.

[FE_Geomitrini-5] 1 2 3 4 5 6 7 8 9 10 11 12 "Geomitrini". Fauna Europaea, last update 27 January 2011, accessed 4 August 2011.

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Anatomy

Taxonomy

References