// Workers AI · dad joke modeWhat did the Escucha Formation say? "I'm a rock solid listener".
| Escucha Formation | |
|---|---|
| Stratigraphic range: Late Aptian-mid Albian ~ | |
.JPG) View of Escucha and surrounding rocks | |
| Type | Geological formation |
| Sub-units | E1 to E3 |
| Underlies | Utrillas Formation |
| Overlies | Castrillo de la Reina, Benassal and Oliete Formations |
| Thickness | ~70 m (230 ft) |
| Lithology | |
| Primary | Siltstone, mudstone (E1, E3) sandstone (E2) |
| Other | Coal (E1) siltstone (E2), amber |
| Location | |
| Coordinates | 42°36′N 2°42′W / 42.6°N 2.7°W |
| Approximate paleocoordinates | 30°24′N 1°24′E / 30.4°N 1.4°E |
| Region | Teruel, Aragón La Rioja |
| Country | Spain |
| Extent | Cameros, Maestrazgo and Oliete Basins |
| Type section | |
| Named for | Escucha, Teruel |
  Escucha Formation (Spain) | |
The Escucha Formation is a geological formation in La Rioja and Teruel provinces of northeastern Spain whose strata date back to the late Aptian to middle Albian stages of the Early Cretaceous. Dinosaur remains are among the fossils that have been recovered from the formation.[1]
The approximately 70 m (230 ft) thick formation underlies the Utrillas Formation and overlies Castrillo de la Reina, Benassal & Oliete Formations. The Escucha Formation comprises siltstones, mudstones, sandstones, coal, siltstones and amber, in which diverse fossil insects were found. The formation was deposited in a variety of continental to paralic (deltaic) environments.[2]
Depositional Environment
[edit]The Escucha Formation represents syn-rift depositional environment, deposited in siliciclastic carbonate platforms, coastal siliciclastic environments and other offshore settings. In the basal part of the formation, cobble to boulder-sized dropstones of glacial origin have been identified, representing long-distance iceberg rafting from Arctic sources.[3]
Fossil content
[edit]The Escucha Formation has provided the following fossils, among others:[4]
Dinosaurs
[edit]| Dinosaurs of the Escucha Formation | ||||||
|---|---|---|---|---|---|---|
| Genus | Species | Region | Member | Material | Notes | Image |
| Proa | P. valdearinnoensis[5] | Santa Maria Mine | Lower Sedimentary Succession Member | Three partial skeletons and several skull elements from different individuals | Hadrosauriform ornithopod |  |
| Europelta | E. carbonensis[6] | Santa Maria Mine | Lower Sedimentary Succession Member | Two partial skeletons | Struthiosaurine ankylosaur |  |
| Allosauroidea indet. | indeterminate[7] | Santa Maria Mine | Lower Sedimentary Succession Member | Tooth | ||
| Titanosauriformes indet.[8] | indeterminate | Partial unla | ||||
Pseudosuchia
[edit]| Pseudosuchians of the Escucha Formation[9][10] | ||||||
|---|---|---|---|---|---|---|
| Genus | Species | Region | Member | Material | Notes | Image |
| Anteophthalmosuchus | A. escuchae | Santa Maria Mine | Lower Sedimentary Succession Member | Partial skeleton including skull, and other isolated remains | Goniopholididae. Partial skull AR-1-3422 represents juvenile individual of this taxon. | |
| Hulkepholis | H. plotos[9] | Santa Maria Mine | Lower Sedimentary Succession Member | Partial skeleton including skull | Goniopholididae | |
Testudines
[edit]| Testudines of the Escucha Formation | ||||||
|---|---|---|---|---|---|---|
| Genus | Species | Region | Member | Material | Notes | Image |
| Aragochersis | A. lignitesta[11] | Santa Maria Mine | Lower Sedimentary Succession Member | Two partial skeletons, both including complete skull | Helochelydridae turtle. | |
| Toremys | T. cassiopeia[12] | Santa Maria Mine | Lower Sedimentary Succession Member | Multiple shells both articulated and disarticulated, and several appendicular elements | Pleurosternidae turtle. | |
| Trachydermochelys | T. sp.[13] | Cabezo de las Eras | Lower Member | Helochelydridae turtle. | ||
Fishes
[edit]| Fishes of the Escucha Formation[7] | ||||||
|---|---|---|---|---|---|---|
| Genus | Species | Region | Member | Material | Notes | Image |
| Chondrichthyes indet. | Indeterminate | Santa Maria Mine | Lower Sedimentary Succession Member | |||
| Osteichthyes indet. | Indeterminate | Santa Maria Mine | Lower Sedimentary Succession Member | |||
Molluscs
[edit]| Molluscs of the Escucha Formation[7] | ||||||
|---|---|---|---|---|---|---|
| Genus | Species | Region | Member | Material | Notes | Image |
| Bivalvia indet. | Indeterminate | Santa Maria Mine | Lower Sedimentary Succession Member | |||
| Gastropoda indet. | Indeterminate | Santa Maria Mine | Lower Sedimentary Succession Member | |||
Crustacea
[edit]| Crustaceans of the Escucha Formation | ||||||
|---|---|---|---|---|---|---|
| Genus | Species | Region | Member | Material | Notes | Image |
| Cretagourretia | C. salasi[14] | CV-13 Roadcut | Ctenochelidae decapod | |||
| Joeranina | J. tausi[14] | CV-13 Roadcut | Palaeocorystidae decapod | |||
Insects
[edit]| Insects | ||||
|---|---|---|---|---|
| Taxa | Location/site | Notes | Images | |
| Alavaraphidia | Peñacerrada I amber | [15] |  | |
| Amarantoraphidia | Peñacerrada I amber | [15] |  | |
| Microcostaphron parvus | Peñacerrada I amber | [16] | ||
| Radiophron ibericus | Peñacerrada I amber | [17] | ||
| Styporaphidia? hispanica | Peñacerrada I amber | [15] |  | |
- Other arthropods
- Alavia neli
- Archaelagonops alavensis
- Mesozygiella dunlopi
- Cretogarypinus zaragozai[18]
- Ithioreolpium alavensis[18]
- Alavesiaphis margaritae
- Hispanocader lisae
- Alavametra popovi
- Iberovelia quisquilia
- Glaesivelia pulcherrima
- Gymnopollisthrips maior
- G. minor
- Manicapsocidus enigmaticus
- Archaeatropos alavensis
- Preempheria antiqua
- Empheropsocus arilloi
- E. margineglabrus
- Morazatermes krishnai
- Cantabritermes simplex
- Autrigonoforceps iberica
- Hispanelcana alavensis
- H. arilloi
- H. lopezvallei
- Libanophron sugaar
- Hippocoon basajauni
- Burmaphron iratxoak
- B. jentilak
- B. sorginak
- Tagsmiphron olentzero
- Elasmophron mari
- Ectenobythus iberiensis
- Liztor pilosus
- Cretepyris martini
- Zophepyris alavaensis
- Ampulicomorpha perialla
- Microcostaphron parvus
- Archephedrus stolamissus
- Protorhyssalopsis perrichoti
- Spathiopteryx alavarommopsis
- Serphites lamiak
- Aposerphites angustus
- Archaeromma hispanicum
- Galloromma alavaensis
- Alavaromma orchamum
- Tithonoscelio resinalis
- Bruescelio platycephalus
- Electroteleiopsis hebdomas
- Alavascelio delvallei
- Amissascelio temporarius
- Perimoscelio tyrbastes
- Perimoscelio confector
- Proterosceliopsis masneri
- Juxtascelio interitus
- Iberopria perialla
- Iberoevania roblesi
- Cretevania alonsoi
- Iberomaimetsha nihtmara
- Iberomaimetsha rasnitsyni
- Valaa delclosi
- Megalava truncata
- Eosyntexis parva
- Cretasonoma corinformibus
- Penarhytus tenebris
- Prosolierius parvus
- Cretakarenni hispanicus
- Rhizophtoma longus
- Darwinylus marcosi
- Mediumiuga sinespinis
- Archiaustroconops alavensis
- Leptoconops (Leptoconops) zherikhini
- Gerontodacus skalskii
- Chimeromyia alava
- Chimeromyina concilia
- Euliphora grimaldii
- Alavesia subiasi
- Tethepomima holomma
- Tethepomyia buruhandi
- Lysistrata emerita
- Hegalari minor
- Hegalari antzinako
- Alavamanota hispanica
- Allocotocera xavieri
- Cretohaplusia ortunoi
- Eltxo cretaceus
- Helius (Helius) alavensis
- Helius (Helius) spiralis
- Espanoderus barbarae
- Alavaraphidia imperterrita
- Baissoptera cretaceoelectra
- Cretokatianna bucculenta
- Sphyrotheciscus senectus
- Archeallacma dolichopoda
- Katiannasminthurus xenopygus
- Pseudosminthurides stoechus
- Burmisotoma spinulifera
- Protoisotoma autrigoniensis
- Proisotoma communis
- Proleptochelia tenuissima
- Electrotanais monolithus
- Alavatanais carabe
- Alavatanais margulisae
- Eurotanais terminator
- Spinomegops arcanus
- Strieremaeus minguezae
- Iberofoveopsis miguelesi
- Hispanothrips utrillensis
- Aragonitermes teruelensis
- Aragonimantis aenigma
- Mymaropsis turolensis
- Serphites silban
- Galloromma turolensis
- Alavaromma orchamum
- Cretevania rubusensis
- Cretevania alcalai
- Cretevania montoyai
- Actenobius magneoculus
- Arra legalovi
- Leptoconops (Leptoconops) zherikhini
- Atriculicoides sanjusti
- Atriculicoides hispanicus
- Gerontodacus skalskii
- Microphorites utrillensis
- Litoleptis fossilis
- Burmazelmira grimaldii
- Aragomantispa lacerata
- Spinomegops aragonensis
- Ametroproctus valeriae
- Cretaceobodes martinezae
- Trhypochthonius lopezvallei
- Invertebrates
- Flora
Correlation
[edit]
See also
[edit]- List of dinosaur-bearing rock formations
- Monte Grande Formation, Albian formation of the Cantabrian Basin
- Caranguejeira Conglomerate, Aptian to Cenomanian formation of the Lusitanian Basin
- Baltic, Burmese, Dominican and Mexican amber
References
[edit]- ↑ Weishampel, et al., 2004
- ↑ Peyrot et al., 2007
- ↑ Rodríguez-López, Juan Pedro; Liesa, Carlos L.; Pardo, Gonzalo; Meléndez, Nieves; Soria, Ana R.; Skilling, Ian (2016-06-15). "Glacial dropstones in the western Tethys during the late Aptian–early Albian cold snap: Palaeoclimate and palaeogeographic implications for the mid-Cretaceous". Palaeogeography, Palaeoclimatology, Palaeoecology. 452: 11–27. Bibcode:2016PPP...452...11R. doi:10.1016/j.palaeo.2016.04.004. ISSN 0031-0182.
- ↑ Escucha Formation at Fossilworks.org
- ↑ Mcdonald, Andrew T.; EspíLez, Eduardo; Mampel, Luis; Kirkland, James I.; Alcalá, Luis (21 December 2012). "An unusual new basal iguanodont (Dinosauria: Ornithopoda) from the Lower Cretaceous of Teruel, Spain". Zootaxa. 3595 (1). Bibcode:2012Zoot.35955.1.3M. doi:10.11646/zootaxa.3595.1.3.
- ↑ Kirkland, James I.; Alcalá, Luis; Loewen, Mark A.; Espílez, Eduardo; Mampel, Luis; Wiersma, Jelle P. (2 December 2013). "The Basal Nodosaurid Ankylosaur Europelta carbonensis n. gen., n. sp. from the Lower Cretaceous (Lower Albian) Escucha Formation of Northeastern Spain". PLOS ONE. 8 (12) e80405. Bibcode:2013PLoSO...880405K. doi:10.1371/journal.pone.0080405. PMC 3847141. PMID 24312471.
- 1 2 3 Alcalá, Luis; Espílez, Eduardo; Mampel, Luis; Kirkland, James I.; Ortiga, Manuel; Rubio, Diego; González, Ana; Ayala, Daniel; Cobos, Alberto; Royo-Torres, Rafael; Gascó, Francisco; Pesquero, Maria Dolores (1 December 2012). "A New Lower Cretaceous Vertebrate Bonebed Near Ariño (Teruel, Aragón, Spain); Found and Managed in a Joint Collaboration Between a Mining Company and a Palaeontological Park". Geoheritage. 4 (4): 275–286. Bibcode:2012Geohe...4..275A. doi:10.1007/s12371-012-0068-y.
- ↑ Canudo, J.I.; Ruiz-Omeñaca, J.I.; Sender, L.M. (2004). "Primera evidencia de un dinosaurio saurópodo en la Formación Escucha (Utrillas, Teruel), Albiense medio (Cretácico inferior)" (PDF). Geo-Temas. 6 (5): 27–30.
- 1 2 Buscalioni, Ángela D; Alcalá, Luis; Espílez, Eduardo; Mampel, Luis (2013). "European Goniopholididae from the Early Albian Escucha Formation in Ariño (Teruel, Aragón, Spain)". Spanish Journal of Palaeontology. 28 (1): 103–122. doi:10.7203/sjp.28.1.17835.
- ↑ Arribas, Ignacio; Buscalioni, Angela D.; Royo Torres, Rafael; Espílez, Eduardo; Mampel, Luis; Alcalá, Luis (31 October 2019). "A new goniopholidid crocodyliform, Hulkepholis rori sp. nov. from the Camarillas Formation (early Barremian) in Galve, Spain)". PeerJ. 7 e7911. Bibcode:2019PeerJ...7e7911A. doi:10.7717/peerj.7911. PMC 6825746. PMID 31687271.
- ↑ Pérez-García, A.; Espílez, E.; Mampel, L.; Alcalá, L. (March 2020). "A new basal turtle represented by the two most complete skeletons of Helochelydridae in Europe". Cretaceous Research. 107 104291. Bibcode:2020CrRes.10704291P. doi:10.1016/j.cretres.2019.104291.
- ↑ Pérez-García, A.; Espílez, E.; Mampel, L.; Alcalá, L. (July 2015). "A new European Albian turtle that extends the known stratigraphic range of the Pleurosternidae (Paracryptodira)". Cretaceous Research. 55: 74–83. Bibcode:2015CrRes..55...74P. doi:10.1016/j.cretres.2015.02.007.
- ↑ Canudo, J. I.; Cobos, A.; Martín-Closas, C.; Murelaga, X.; Pereda-Suberbiola, X.; Royo-Torres, R.; Ruiz-Omeñaca, J. I.; Sender, L. M. (2005). "Sobre la presencia de dinosaurios ornitópodos en la Formación Escucha (Cretácico Inferior, Albiense): redescubierto "Iguanodon" en Utrillas (Teruel)". XVI Reunión Bienal de la Real Sociedad Española de Historia Natural, Fundación Conjunto Paleontológico de Teruel, Teruel, Spain. Fundamental 6: 51–56.
- 1 2 Ossó, Àlex; Charbonnier, Sylvain; Hyžný, Matúš; Van Bakel, Barry W.M.; Devillez, Julien; Bover-Arnal, Telm; Moreno-Bedmar, Josep Anton (24 October 2024). "Decapod assemblages from the Aptian-Albian transition of the eastern Maestrat Basin (Iberian Chain)". Geologica Acta. 22: 1–35. Bibcode:2024GeolA..22....7O. doi:10.1344/GeologicaActa2024.22.7.
- 1 2 3 Pérez de la Fuente et al., 2012
- ↑ Ortega Blanco et al., 2010, p.272
- ↑ Ortega Blanco et al., 2010, p.267
- 1 2 Sánchez-García, A.; Palencia, L.; Delclòs, X.; Peñalver, E. (2024). "False scorpions (Arachnida, Pseudoscorpiones) from Lower Cretaceous Spanish amber". Papers in Palaeontology. 10 (6). e1608. Bibcode:2024PPal...10E1608S. doi:10.1002/spp2.1608. hdl:10550/111887.
Bibliography
[edit]- Kirkland, J.I.; Alcalá, L.; Loewen, M.A.; Espílez, E.; Mampel, L.; Wiersma, J.P. (2013). "The Basal Nodosaurid Ankylosaur Europelta carbonensis n. gen., n. sp. From the Lower Cretaceous (Lower Albian) Escucha Formation of Northeastern Spain". PLoS ONE. 8 (12) e80405. Bibcode:2013PLoSO...880405K. doi:10.1371/journal.pone.0080405. PMC 3847141. PMID 24312471.
- Ortega Blanco, J.; Rasnitsyn, A.P.; Delclòs, X. (2010). "A new family of ceraphronoid wasps from Early Cretaceous Álava Amber, Spain". Acta Palaeontologica Polonica. 55 (2): 265–276. Bibcode:2010AcPaP..55..265O. doi:10.4202/app.2009.0014. S2CID 58934033. Retrieved 2019-10-09.
- Pérez de la Fuente, R.; Peñalver, E.; Delclòs, X.; Engel, M.S. (2012). "Snakefly diversity in Early Cretaceous amber from Spain (Neuropterida, Raphidioptera)". ZooKeys (204): 1–40. Bibcode:2012ZooK..204....1P. doi:10.3897/zookeys.204.2740. PMC 3391719. PMID 22787417.
- Pérez García, A.; Espílez, E.; Mampel, L.; Alcalá, L. (2020). "A new basal turtle represented by the two most complete skeletons of Helochelydridae in Europe". Cretaceous Research. 107 104291. Bibcode:2020CrRes.10704291P. doi:10.1016/j.cretres.2019.104291. S2CID 210279023.
- Peyrot, Daniel; Rodríguez López, Juan Pedro; Lassaletta, Luis; Meléndez, Nieves; Barrón, Eduardo (2007). "Contributions to the palaeoenvironmental knowledge of the Escucha Formation in the Lower Cretaceous Oliete Sub-basin, Teruel, Spain". Comptes Rendus Palevol. 6 (6–7): 469–481. Bibcode:2007CRPal...6..469P. doi:10.1016/j.crpv.2007.09.014.
- Weishampel, David B.; Dodson, Peter; Osmólska, Halszka, eds. (2004). The Dinosauria, 2nd edition. Berkeley: University of California Press. pp. 1–880. ISBN 978-0-520-94143-4. Retrieved 2019-02-21.
{{cite book}}: CS1 maint: publisher location (link)
Further reading
[edit]- L. Alcalá, E. Espílez, L. Mampel, J. I. Kirkland, M. Ortiga, D. Rubio, A. González, D. Ayala, A. Cobos, R. Royo-Torres, F. Gascó and M. D. Pesquero. 2012. A new Lower Cretaceous vertebrate bonebed near Arino (Tereul, Aragon, Spain); found and managed in a joint collaboration between a mining company and a palaeontological park. Geoheritage 4:275-286
- J. I. Canudo, A. Cobos, C. Martín-Closas, X. Murelaga, X. Pereda-Suberbiola, R. Royo-Torres, J. I. Ruiz-Omeñaca and L. M. Sender. 2005. Sobre la presencia de dinosaurios ornitópodos en la Formación Escucha (Cretácico Inferior, Albiense): redescubierto "Iguanodon" en Utrillas (Teruel) [On the presence of ornithopod dinosaurs in the Escucha Formation (Lower Cretaceous, Albian): redescribing "Iguanodon" in Utrillas (Teruel)]. Fundamental 6:51-56
- E. Peñalver, D. A. Grimaldi, and X. Delclos. 2006. Early Cretaceous spider web with its prey. Science 312:1761