Udurchukan Formation

Udurchukan Formation
Udurchukan Formation
Stratigraphic range: Maastrichtian; ~70–66 Ma PreꞒ Ꞓ O S D C P T J K Pg N
Type	Geological formation
Unit of	Tsagayan Group
Underlies	Bureya Formation
Overlies	Kundur Formation
Thickness	11 metres (40 ft) exposed at Kundur locality
Lithology
Primary	Conglomerate, mudstone, diamictite
Other	Sandstone
Location
Coordinates	49°30′N 129°30′E / 49.5°N 129.5°E
Approximate paleocoordinates	51°24′N 115°48′E / 51.4°N 115.8°E
Region	Amur Oblast
Country	Russia
Extent	Zeya-Bureya Basin
	Udurchukan Formation (Russia) Udurchukan Formation (Amur Oblast)

The Udurchukan Formation is a geological formation located in Amur Region, Far East Russia. Based on palynomorphs such as Wodehouseia spinata, the Udurchukan is considered of Maastrichtian age of the Late Cretaceous, during the Cretaceous Period.

Fossil record

Since Wodehouseia spinata and Aquillapollenites subtilis are known in the Americas only from the Late Maastrichtian, the presence of these palynomorphs in the Udurchukan caused Godefroit to consider the unit and its lambeosaurine dominated fauna to be coeval with the Lance Formation and Hell Creek Formation. However, research in the Songliao Basin indicates Wodehouseia spinata is also known from the early (albeit not basal) and middle Maastrichtian of Asia.

Dinosaurs

Color key

Taxon	Reclassified taxon	Taxon falsely reported as present	Dubious taxon or junior synonym	Ichnotaxon	Ootaxon	Morphotaxon

Notes
Uncertain or tentative taxa are in small text; ~~crossed out~~ taxa are discredited.

Ornithischians

Ankylosaurs

Ankylosaurs reported from the Udurchukan Formation
Taxon	Species	Location	Stratigraphic position	Material	Notes	Images
Nodosauridae^[1]	indet.	Kundur	Upper	A tooth, an incomplete tooth and an osteo dermal scute.	An indeterminate nodosauridae.

Hadrosaurs

Hadrosaurs reported from the Udurchukan Formation
Taxon	Species	Location	Stratigraphic position	Material	Notes	Images
Amurosaurus	A. riabinini	Blagoveschensk		A partial remnants of skull and lower jaw.	A lambeosaurine hadrosaur which was the most common dinosaur in the area, a bonebed containing many specimens was unearthed in 2008.^[2]
Kerberosaurus^[3]	K. manakini	Blagoveschensk		The caudal part of cranium consists of a braincase.	A saurolophine hadrosaur
Kundurosaurus^[4]	K. nagornyi	Kundur		A partial, disarticulated skull.	A saurolophine hadrosaur; possibly synonymous with Kerberosaurus
Hadrosaurinae indet.	Indetermidate
Olorotitan^[5]	O. arharensis	Kundur		A nearly complete skeleton.	A lambeosaurine hadrosaur

Sauropods

Sauropods reported from the Udurchukan Formation
Taxon	Species	Location	Stratigraphic position	Material	Notes	Images
Arkharavia^[6]	A. heterocoelica	Kundur	Upper	A single tooth and a proximal tail vertebrae, although some remains probably belong to a hadrosaurid.	A somphospondylan sauropod.^[7]
Opisthocoelicaudiinae indet.^[8]	Indetermidate
Titanosauria indet.	Indetermidate

Theropods

Theropods reported from the Udurchukan Formation
Taxon	Species	Location	Stratigraphic position	Material	Notes	Images
Albertosaurinae indet.	Indetermidate
Dromaeosaurinae indet.	Indetermidate
Dromaeosaurinae indet.	Indeterminate
Ornithomimidae indet.^[9]	Indetermidate				Described as being very similar to Qiupalong^[9]
Richardoestesia	R. sp
Saurornitholestes	S. cf. sp.
Theropoda indet.	Indeterminate
Troodon	T. sp.
Tyrannosaurinae indet.	Indetermidate				Similar in size to Tyrannosaurus^[10]

Reptiles

Reptiles reported from the Udurchukan Formation
Taxon	Species	Location	Stratigraphic position	Material	Notes	Images
Amuremys	A. planicostata
Crocodylia indet.	Indeterminate
Lindholmemydidae indet.	Indeterminate
Shamosuchus	S. sp.
Testudines indet.	Indeterminate
Trionychidae indet.	Indeterminate

Mammals

Mammals reported from the Udurchukan Formaton
Taxon	Species	Location	Stratigraphic Position	Material	Notes	Images
Cimolodonta indet.	Indeterminate

Insects

Insects reported from the Udurchukan Formation
Taxon	Species	Location	Stratigraphic position	Material	Notes	Images
Mesosigara	M. kryshtofovichi

Flora

Plants reported from the Udurchukan Formation
Taxon	Species	Location	Stratigraphic position	Material	Notes	Images
Cupressinoxylon	C. sp.

References

↑ Tumanova, Tatiana & Bolotsky, Yu.L. & Alifanov, Vladimir. (2004). The first finds of armored dinosaurs in the Upper Cretaceous of Russia (Amur Region). Paleontological Journal. 38. 73-77.
↑ Lauters, Pascaline; Bolotsky, Yuri L.; Van Itterbeeck, Jimmy; Godefroit, Pascal (March 1, 2008). "Taphonomy and Age Profile of a Latest Cretaceous Dinosaur Bone Bed in Far Eastern Russia". PALAIOS. 23 (3). SEPM Society for Sedimentary Geology: 153–162. Bibcode:2008Palai..23..153L. doi:10.2110/palo.2006.p06-031r. Retrieved 1 May 2021.
↑ Bolotsky, Yu.L.; Godefroit, P. (2004). "A new hadrosaurine dinosaur from the Late Cretaceous of far Eastern Russia". Journal of Vertebrate Paleontology. 24 (2): 351–365. Bibcode:2004JVPal..24..351B. doi:10.1671/1110. S2CID 130691286.
↑ Pascal Godefroit; Yuri L. Bolotsky; Pascaline Lauters (2012). "A New Saurolophine Dinosaur from the Latest Cretaceous of Far Eastern Russia". PLOS ONE. 7 (5) e36849. Bibcode:2012PLoSO...736849G. doi:10.1371/journal.pone.0036849. PMC 3364265. PMID 22666331.
↑ Godefroit, Pascal; Bolotsky, Yuri; Alifanov, Vladimir (2003). "A remarkable hollow-crested hadrosaur from Russia: an Asian origin for lambeosaurines" (PDF). Comptes Rendus Palevol. 2 (2). Cellule MathDoc/CEDRAM: 143–151. Bibcode:2003CRPal...2..143G. doi:10.1016/s1631-0683(03)00017-4. ISSN 1631-0683.
↑ Alifanov, Vladimir & Bolotsky, Yu. (2010). Arkharavia heterocoelica gen. et sp. nov., a new sauropod dinosaur from the Upper Cretaceous of the Far East of Russia. Paleontological Journal. 44. 84-91. 10.1134/S0031030110010119.
↑ Mannion, Philip D.; Upchurch, Paul; Barnes, Rosie N.; Mateus, Octávio (2013). "Osteology of the Late Jurassic Portuguese sauropod dinosaur Lusotitan atalaiensis (Macronaria) and the evolutionary history of basal titanosauriforms". Zoological Journal of the Linnean Society. 168: 98–206. doi:10.1111/zoj.12029.
↑ Averianov, A.O.; Bolotsky, YL; Bolotsky, YI (2023-03-14). "A Sauropod from the Late Cretaceous of the Amur Region". Doklady Earth Sciences. 507: S418–S419. doi:10.1134/S1028334X22601547. S2CID 257498576.
1 2 Averianov, Alexander; Skutschas, Pavel; Bolotsky, Yuriy; Bolotsky, Ivan (2023-12-31). "First find of an ornithomimid theropod dinosaur in the Upper Cretaceous of the Russian Far East". Biological Communications. 68 (4). doi:10.21638/spbu03.2023.405. ISSN 2587-5779.
↑ Yun, Changyu. "Testing the hypotheses of the origin of Tyrannosaurus rex: immigrant species, or native species?" (PDF). The Journal of Paleontological Sciences: 1–9.

Bibliography

Y. A. Popov. 1971. Historical development of the infraorder Nepomorpha (Heteroptera). Akademiya Nauk SSSR, Trudy Paleontologicheskogo Instituta 129:1-228

[1] Tumanova, Tatiana & Bolotsky, Yu.L. & Alifanov, Vladimir. (2004). The first finds of armored dinosaurs in the Upper Cretaceous of Russia (Amur Region). Paleontological Journal. 38. 73-77.

[2] Lauters, Pascaline; Bolotsky, Yuri L.; Van Itterbeeck, Jimmy; Godefroit, Pascal (March 1, 2008). "Taphonomy and Age Profile of a Latest Cretaceous Dinosaur Bone Bed in Far Eastern Russia". PALAIOS. 23 (3). SEPM Society for Sedimentary Geology: 153–162. Bibcode:2008Palai..23..153L. doi:10.2110/palo.2006.p06-031r. Retrieved 1 May 2021.

[3] Bolotsky, Yu.L.; Godefroit, P. (2004). "A new hadrosaurine dinosaur from the Late Cretaceous of far Eastern Russia". Journal of Vertebrate Paleontology. 24 (2): 351–365. Bibcode:2004JVPal..24..351B. doi:10.1671/1110. S2CID 130691286.

[4] Pascal Godefroit; Yuri L. Bolotsky; Pascaline Lauters (2012). "A New Saurolophine Dinosaur from the Latest Cretaceous of Far Eastern Russia". PLOS ONE. 7 (5) e36849. Bibcode:2012PLoSO...736849G. doi:10.1371/journal.pone.0036849. PMC 3364265. PMID 22666331.

[5] Godefroit, Pascal; Bolotsky, Yuri; Alifanov, Vladimir (2003). "A remarkable hollow-crested hadrosaur from Russia: an Asian origin for lambeosaurines" (PDF). Comptes Rendus Palevol. 2 (2). Cellule MathDoc/CEDRAM: 143–151. Bibcode:2003CRPal...2..143G. doi:10.1016/s1631-0683(03)00017-4. ISSN 1631-0683.

[6] Alifanov, Vladimir & Bolotsky, Yu. (2010). Arkharavia heterocoelica gen. et sp. nov., a new sauropod dinosaur from the Upper Cretaceous of the Far East of Russia. Paleontological Journal. 44. 84-91. 10.1134/S0031030110010119.

[mannionetal-7] Mannion, Philip D.; Upchurch, Paul; Barnes, Rosie N.; Mateus, Octávio (2013). "Osteology of the Late Jurassic Portuguese sauropod dinosaur Lusotitan atalaiensis (Macronaria) and the evolutionary history of basal titanosauriforms". Zoological Journal of the Linnean Society. 168: 98–206. doi:10.1111/zoj.12029.

[8] Averianov, A.O.; Bolotsky, YL; Bolotsky, YI (2023-03-14). "A Sauropod from the Late Cretaceous of the Amur Region". Doklady Earth Sciences. 507: S418–S419. doi:10.1134/S1028334X22601547. S2CID 257498576.

[ornithomimidaeindet-9] 1 2 Averianov, Alexander; Skutschas, Pavel; Bolotsky, Yuriy; Bolotsky, Ivan (2023-12-31). "First find of an ornithomimid theropod dinosaur in the Upper Cretaceous of the Russian Far East". Biological Communications. 68 (4). doi:10.21638/spbu03.2023.405. ISSN 2587-5779.

[10] Yun, Changyu. "Testing the hypotheses of the origin of Tyrannosaurus rex: immigrant species, or native species?" (PDF). The Journal of Paleontological Sciences: 1–9.

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