Batylykh Formation

Batylykh Formation
Batylykh Formation
Stratigraphic range: Early Cretaceous; ?Berriasian–Barremian PreꞒ Ꞓ O S D C P T J K Pg N
	Teete locality with stratigraphic column
Type	Geological formation
Unit of	Sangar Series
Underlies	Eksenyakh Formation
Lithology
Primary	Claystone, siltstone, sandstone
Location
Coordinates	62°42′N 117°36′E / 62.7°N 117.6°E
Approximate paleocoordinates	66°30′N 110°06′E / 66.5°N 110.1°E
Region	Yakutia
Country	Russia
Extent	Vilyuy River Basin
	Batylykh Formation (Russia)

The Batylykh Formation is a geological formation in Yakutia, Russia. It is of an uncertain Early Cretaceous age, probably dating between the Berriasian and the Barremian. It is the oldest unit of the 1,500 metres (4,900 ft) thick Sangar Series within the Vilyuy syneclise. The mudstones, sandstones and shales of the formation were deposited in a fluvial to lacustrine environment.

Fossil content

The formation is best known for the Teete locality, which has revealed remains of numerous vertebrates, including dinosaurs and mammals.

Fossils recovered from the formation include:^[1]

Color key

Taxon	Reclassified taxon	Taxon falsely reported as present	Dubious taxon or junior synonym	Ichnotaxon	Ootaxon	Morphotaxon

Notes
Uncertain or tentative taxa are in small text; ~~crossed out~~ taxa are discredited.

Dinosaurs

Ornithischians

Ankylosaurs

Ankylosaurs of the Batylykh Formation
Genus	Species	Location	Stratigraphic position	Material	Notes	Image
Polacanthinae Indet.	Indeterminate	Sakha Republic, Russia	Berriasian to Barremian		A polacanthine nodosaurid

Stegosaurs

Stegosaurs of the Batylykh Formation
Genus	Species	Location	Stratigraphic position	Material	Notes	Image
Stegosaurinae Indet.^[2]	Indeterminate	Sakha Republic, Russia	Berriasian to Barremian		A stegosaurine stegosaurid

Saurischians

Sauropods

Sauropods of the Batylykh Formation
Genus	Species	Location	Stratigraphic position	Material	Notes	Image
Euhelopodidae Indet.	Indeterminate	Sakha Republic, Russia	Berriasian to Barremian		A euhelopodid somphospondylian
Titanosauriformes Indet.	Indeterminate	Sakha Republic, Russia	Berriasian to Barremian		A titanosauriform sauropod

Theropods

Theropods of the Batylykh Formation
Genus	Species	Location	Stratigraphic position	Material	Notes	Image
Compsognathidae Indet.	Indeteminate	Sakha Republic, Russia	Berriasian–Barremian		A compsognathid theropod
Proceratosauridae Indet.	Indeterminate	Sakha Republic, Russia	Berriasian to Barremian		A proceratosaurid tyrannosauroid

Crocodylomorphs

Crocodylomorphs of the Batylykh Formation
Genus	Species	Location	Stratigraphic position	Material	Notes	Image
Goniopholididae Indet.	Indeterminate	Sakha Republic, Russia	Berriasian to Barremian		A goniopholidid neosuchian

Choristoderans

Choristoderans of the Batylykh Formation
Genus	Species	Location	Stratigraphic position	Material	Notes	Image
Khurendukhosaurus	K. sp.	Sakha Republic, Russia	Berriasian to Barremian		A choristoderan

Squamates

Squamates of the Batylykh Formation
Genus	Species	Location	Stratigraphic position	Material	Notes	Image
Scincomorpha Indet.	Indeterminate	Sahka Republic, Russia	Berriasian to Barremian		A scincomorph lizard

Turtles

Turtles of the Batylykh Formation
Genus	Species	Location	Stratigraphic position	Material	Notes	Image
Helochelydridae Indet.^[3]	Indeterminate	Sakha Republic, Russia	Berriasian to Barremian		A helochelydrid turtle

Amphibians

Amphibians of Batylykh Formation
Genus	Species	Location	Stratigraphic position	Material	Notes	Image
Karauridae Indet.^[4]	Indeterminate	Sakha Republic, Rusia	Berriasian to Barremian		A stem-group salamander
Kulgeriherpeton^[5]	K. ultimus	Sakha Republic, Russia	Berriaman to Barremian		An early salamander
Pelobatidae Indet.	Indeterminate	Sakha Republic, Russia	Berriasian to Barremian		A pelobatid frog

Fish

Fishes of the Batylykh Formation
Genus	Species	Location	Stratigraphic position	Material	Notes	Image
Palaeoniscidae Indet.	Indeterminate	Sakha Republic, Russia	Berriasian to Barremian		palaeoniscid fish
Peipiaosteidae Indet.	Indeterminate	Sakha Republic, Russia			A peipiaosteid acipenseriform

Mammals

Mammals of the Batylykh Formation
Genus	Species	Location	Stratigraphic position	Material	Notes	Image
Cryoharamiya^[6]	C. tarda	Sakha Republic, Russia	Berriasian to Barremian		A euharamiyidan mammaliaform
Docodonta Indet.	Indeterminate	Sakha Republic, Russia	Berriasian to Barremian		An docodont
Gobiconodon	G. sp.	Sakha Republic, Russia	Berriasian to Barremian		A gobinonodontid eutriconodont
Khorotherium	K. yakutensis	Sakha Republic, Russia	Berriasian to Barremian		tegotheriid docodont
Sangarotherium^[7]	S. aquilonium	Sakha Republic, Russia	Berriasan to Barremian		An eutriconodont
cf. Sineleutherus	cf. S. sp.	Sakha Republic, Russia	Berriasian to Barremian		eleutherodontid euharamiyidan
Tritylodontidae Indet.^[8]	Indeterminate	Sakha Republic, Russia	Berriasian to Barremian		A tritylodontid cynodont
Xenocretosuchus	X. kolossovi	Sakha Republic, Russia	Berriasian to Barremian		A tritylodontid cynodont

Insects

Insects of the Batylykh Formation
Genus	Species	Location	Stratigraphic position	Material	Notes	Image
Crenoptychoptera^[9]	C. gronskayae	Sakha Republic, Russia	Berriasian to Barremian
Zhiganka^[9]	Z. comitans	Sakha Republic, Russia	Berriasian to Barremian

Mollusks

Bivalves

Bivalves of the Batylykh Formation
Genus	Species	Location	Stratigraphic position	Material	Notes	Image
Bivalvia Indet.	Indeterminate	Sakha Republic, Russia	Berriasian to Barremian

Gastropods

Ankylosaurs of the Grès de Labarre
Genus	Species	Location	Stratigraphic position	Material	Notes	Image
Gastropoda Indet.

Plants

Plants of the Batylykh Formation
Genus	Species	Location	Stratigraphic position	Material	Notes	Image
Ginkgocycadophytus	G. sp.	Sakha Republic, Russia	Berriasian to Barremian
Klukisporites	K. sp.	Sakha Republic, Russia	Berriasian to Barremian
Leiotriletes	L. sp.	Sakha Republic, Russia	Berriasian to Barremian
Ophioglossum	O. sp.	Sakha Republic, Russia	Berriasian to Barremian
Picea	P. sp.	Sakha Republic, Russia	Berriasian to Barremian
Pinaceae Indet.	Indeterminate	Sakha Republic, Russia	Berriasian to Barremian
Piceapollenites	P. sp.	Sakha Republica	Berriasian to Barremian
Pinus	P. sp.	Sakha Republic, Russia	Berriasian to Barremian
Podocarpus	P. multesima	Sakha Republic, Russia	Berriasian to Barremian
Podocarpus	P. sp.	Sakha Republic, Russia	Berriasian to Barremian
Selaginella	S. velata	Sakha Republic, Russia	Berriasian to Barremian
Selaginella	S. sp.	Sakha Republic, Russia	Berriasian to Barremian
Stereisporites	S. compactus	Sakha Republic, Russia	Berriasian to Barremian
	S. congregatus
	S. sp.
Tripartina	T. variabilis	Sakha Republic, Russia	Berriasian to Barremian

Gallery

Dinosaur teeth
Kulgeriherpeton ultimum

References

↑ Batylykh Formation at Fossilworks.org
↑ Skutschas PP, Gvozdkova VA, Averianov AO, Lopatin AV, Martin T, Schellhorn R, et al. (2021). "Wear patterns and dental functioning in an Early Cretaceous stegosaur from Yakutia, Eastern Russia". PLOS ONE. 16 (3) e0248163. Bibcode:2021PLoSO..1648163S. doi:10.1371/journal.pone.0248163. PMC 7968641. PMID 33730093.
↑ Skutschas et al., 2020
↑ Skutschas, Pavel P.; Kolchanov, Veniamin V.; Averianov, Alexander O.; Schellhorn, Rico; Kolosov, Petr N.; Jones, Marc E.H.; Martin, Thomas (August 2023). "The northernmost occurrence of non-karaurid salamanders (Lissamphibia, Caudata) in the Mesozoic". Cretaceous Research. 152 105686. Bibcode:2023CrRes.15205686S. doi:10.1016/j.cretres.2023.105686.
↑ Skutschas et al., 2018
↑ Averianov et al., 2020
↑ Averianov, Alexander O.; Martin, Thomas; Lopatin, Alexey V.; Skutschas, Pavel P.; Vitenko, Dmitry D.; Schellhorn, Rico; Kolosov, Petr N. (August 2023). "On the way from Asia to America: eutriconodontan mammals from the Early Cretaceous of Yakutia, Russia". The Science of Nature. 110 (4): 40. Bibcode:2023SciNa.110...40A. doi:10.1007/s00114-023-01868-3. ISSN 0028-1042. PMID 37530873.
↑ Lopatin & Agadjanian, 2008
1 2 Lukashevich, 1995

Bibliography

Averianov, Alexander O.; Martin, Thomas; Lopatin, Alexey V.; Skutschas, Pavel P.; Schellhorn, Rico; Kolosov, Petr N.; Vitenko, Dmitry D. (2020), "A new euharamiyidan mammaliaform from the Lower Cretaceous of Yakutia, Russia", Journal of Vertebrate Paleontology, 39 (6) e1762089, doi:10.1080/02724634.2019.1762089
Skutschas, P.P.; Markova, V.D.; Kolchanov, V.V.; Averianov, A.O.; Martin, T.; Schellhorn, R.; Kolosov, P.N.; Grigoriev, D.V.; Vitenko, D.D., E. Mobraztsova and I. G. Danilov (2020), "Basal turtle material from the Lower Cretaceous of Yakutia (Russia) filling the gap in the Asian record", Cretaceous Research, x: 1–14, Bibcode:2020CrRes.10604186S, doi:10.1016/j.cretres.2019.07.016{{citation}}: CS1 maint: multiple names: authors list (link)
Averianov, Alexander O.; Skutschas, Pavel P.; Schellhorn, Rico; Lopatin, Alexey V.; Kolosov, Petr N.; Kolchanov, Veniamin V.; Vitenko, Dmitry D.; Grigoriev, Dmitry V.; Martin, Thomas (2019), "The northernmost sauropod record in the Northern Hemisphere: Polar sauropods", Lethaia, 53 (3): 362–368, doi:10.1111/let.12362
Averianov, Alexander; Martin, Thomas; Lopatin, Alexey; Skutschas, Pavel; Schellhorn, Rico; Kolosov, Petr; Vitenko, Dmitry (2018), Fiorillo, Anthony (ed.), "A high-latitude fauna of mid-Mesozoic mammals from Yakutia, Russia", PLoS ONE, 13 (7) e0199983, Bibcode:2018PLoSO..1399983A, doi:10.1371/journal.pone.0199983, ISSN 1932-6203, PMC 6059412, PMID 30044817
Skutschas, P.P.; Kolchanov, V.V.; Averianov, A.O.; Martin, T.; Schellhorn, R.; Kolosov, P.N.; Vitenko, D.D. (2018), "A new relict stem salamander from the Early Cretaceous of Yakutia, Siberian Russia", Acta Palaeontologica Polonica, 63: 519–525, doi:10.4202/app.00498.2018
Kolosov, P.N.; Ivensen, G.V.; Mikhailova, T.E.; Kurzanov, S.M.; Efimov, M.B.; Gubin, Y.M. (2009), "Taphonomy of the Upper Mesozoic Tetrapod Teete Locality (Yakutia)", Paleontological Journal, 43 (2): 201–207, Bibcode:2009PalJ...43..201K, doi:10.1134/S0031030109020129
Lopatin, Alexey; Agadjanian, A. (2008), "A Tritylodont (Tritylodontidae, Synapsida) from the Mesozoic of Yakutia", Doklady Biological Sciences, 419: 107–110, doi:10.1134/S0012496608020117, PMID 18536275
Lukashevich, E.D (1995), "First pupae of the Eoptychopteridae and Ptychopteridae from the Mesozoic of Siberia (Insecta: Diptera)", Paleontological Journal, 29: 164–171