Pterostichinae

Pterostichinae
Image of the common Eurasian species Poecilus cupreus
Scientific classification
Kingdom:	Animalia
Phylum:	Arthropoda
Clade:	Pancrustacea
Class:	Insecta
Order:	Coleoptera
Suborder:	Adephaga
Family:	Carabidae
Subfamily:	Pterostichinae Bonelli, 1810

Pterostichinae is a subfamily of ground beetles (family Carabidae). It belongs to the advanced harpaline assemblage, and if these are circumscribed sensu lato as a single subfamily, Pterostichinae are downranked to a tribe Pterostichini. However, as the former Pterostichitae supertribe of the Harpalinae as loosely circumscribed does seem to constitute a lineage rather distinct from Harpalus, its core group is here considered to be the present subfamily and the Harpalinae are defined more narrowly.

They are usually mid-sized and rather stout ground beetles. Coloration is typically dark and without conspicuous patterns, but often with a strong sheen like polished metal. They are widely distributed and inhabit a wide range of terrestrial habitats. Unlike the more basal ground beetles which only eat small animals, the Pterostichinae include a large proportion of omnivorous or even herbivorous taxa.

Systematics

This group includes the following tribes and genera:

Tribe Chaetodactylini Tschitscherine, 1903

• Chaetodactyla Tschitscherine, 1897

Tribe Cnemalobini Germain, 1911

• Cnemalobus Guerin-Meneville, 1838

Tribe Cratocerini Lacordaire, 1854

• Abacaelostus Straneo, 1952
• Andrewesinulus Straneo, 1938
• Apsidocnemus Alluaud, 1936
• Barylaus Liebherr, 1985
• Brachidius Chaudoir, 1852
• Caecocaelus Straneo, 1949
• Caelostomus W.S.MacLeay, 1825
• Camptogenys Tschitscherine, 1899
• Capabatus Csiki, 1930
• Catapiesis Solier, 1835
• Cratocerus Dejean, 1829
• Crenulostrigus Straneo, 1942
• Cyrtolaus Bates, 1882
• Dactyleurys Tschitscherine, 1899
• Dactylinius Straneo, 1941
• Diachipteryx Alluaud, 1925
• Diceromerus Chaudoir, 1873
• Dromistomus Jeannel, 1948
• Drymonaxus Straneo, 1941
• Feostoma Straneo, 1941
• Hemitelestus Alluaud, 1895
• Homalomorpha Brullé, 1837
• Hoplizomenus Chaudoir, 1873
• Leleuporites Straneo, 1960
• Madapelmus Dajoz, 1985
• Monodryxus Straneo, 1942
• Oxyglychus Straneo, 1938
• Pachycaecus Straneo, 1971
• Pachyroxochus Straneo, 1942
• Platyxythrius Lorenz, 1998
• Stegazopteryx Will, 2004
• Stomonaxellus Tschitscherine, 1901
• Strigomerodes Straneo, 1939
• Strigomerus Chaudoir, 1873
• Trichillinus Straneo, 1938

Tribe Microcheilini Jeannel, 1948

• Microcheila Brulle, 1834

Tribe Morionini Brulle, 1835

• Buderes Murray, 1857
• Hyperectenus Alluaud, 1935
• Hyperion Castelnau, 1834
• Megamorio Chaudoir, 1880
• Morion Latreille, 1810
• Morionidius Chaudoir, 1880
• Moriosomus Motschulsky, 1864
• Platynodes Westwood, 1846
• Stereostoma Murray, 1857

Tribe Pterostichini Bonelli, 1810

• (See Pterostichini for ~180 genera)

Tribe Zabrini Bonelli, 1810

• (See Zabrini for 3 genera)