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Clade of entelegyn spiders
Marronoid clade is a clade of entelegyn spiders classified within the RTA clade . The name "Marronoid clade" was proposed by Wheeler et al. in 2017. The name comes from the Spanish word for brown(marrón ), emphasizing the brown coloration of most of its members. Families in this clade have been used as wastebin taxons and its hard to point out striking synapomorphies for the clade.[ 1] tarsal claws , most families have members both with and without cribellum .[ 1] [ 2]
Currently, the clade consists of 13 families[ 3] [ 4]
Phylogeny of the Marronoid clade based on Gorneau et al. 2023 with the taxonomical revisions proposed by Montana et al. 2025[ 3] [ 4]
sensu Montana et al. 2025
Phylogeny of the Marronoid clade based on Gorneau et al. 2023[ 3]
Phylogeny of the Marronoid clade based on Kulkarni et al. 2023[ 2]
Marronoid clade
Amaurobiidae I
Cycloctenidae I
Stiphidiidae I
Cycloctenidae II
Amaurobiidae II
Hahniidae I
Cybaeidae I
Hahniidae II & III
Cybaeidae II
Phylogeny of the Marronoid clade based on Wheeler et al. 2017[ 1]
Marronoid Clade
Amaurobiidae: Amaurobiinae
Agelenidae
Amaurobiidae: Macrobuninae
Marronoid clade in the phylogeny of the RTA clade , based on Kulkarni et al. 2023[ 2]
RTA Clade
Sparassidae
Marronoid Clade
Marronoid clade in the phylogeny of the RTA clade , based on Wheeler et al. 2017[ 1]
1 2 3 4 Wheeler, W.C.; Coddington, J.A.; Crowley, L.M.; Dimitrov, D.; Goloboff, P.A.; Griswold, C.E.; Hormiga, G.; Prendini, L.; Ramírez, M.J.; Sierwald, P.; Almeida-Silva, L.; Alvarez-Padilla, F.; Arnedo, M.A.; Benavides Silva, L.R.; Benjamin, S.P. (2017). "The spider tree of life: phylogeny of Araneae based on target-gene analyses from an extensive taxon sampling" . Cladistics . 33 (6): 574– 616. doi :10.1111/cla.12182 . 1 2 3 Kulkarni, S.; Wood, H.M.; Hormiga, G. (2023). "Advances in the reconstruction of the spider tree of life: A roadmap for spider systematics and comparative studies" . Cladistics . 39 (6): 479– 532. doi :10.1111/cla.12557 1 2 3 Gorneau, J.A.; Crews, S.C.; Cala-Riquelme, F.; Montana, K.O.; Spagna, J.C.; Ballarin, F.; Almeida-Silva, L.M.; Esposito, L.A. (2023). "Webs of intrigue: museum genomics elucidate relationships of the marronoid spider clade (Araneae)" . Insect Systematics and Diversity . 7 (5) 5: 1– 18. doi :10.1093/isd/ixad021 . 1 2 Monatana, K.O.; Cala-Riquelme, F.; Crews, S.C.; Gorneau, J.A; Al-Jamal, A.M.; Alequín, L.D.; Spagna, J.C.; Ballarin, F.; Esposito, L.A. (2025). "Tailor's drawer no more: a reappraisal of the spider family Dictynidae O. Pickard-Cambridge, 1871 sensu lato" Zoological Journal of the Linnean Society . 204 (2) zlaf007. doi :10.1093/zoolinnean/zlaf007 .
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