// Workers AI · dad joke modeWhat did the Zopheridae beetle say? "I'm beetle down with work.

Zopheridae; Temporal range: Cenomanian–Recent PreꞒ Ꞓ O S D C P T J K Pg N
	Ironclad beetle, Zopherus nodulosus (Zopherinae)
	Aprostoma planifrons (Colydiinae)
Scientific classification
Kingdom:	Animalia
Phylum:	Arthropoda
Clade:	Pancrustacea
Class:	Insecta
Order:	Coleoptera
Suborder:	Polyphaga
Infraorder:	Cucujiformia
Superfamily:	Tenebrionoidea
Family:	Zopheridae; Solier, 1834
Subfamilies
	Colydiinae - cylindrical bark beetles; Zopherinae - ironclad beetles

Zopheridae is a family of beetles belonging to Tenebrionoidea. It has grown considerably in recent years as the members of two other families are now included (Monommatidae and Colydiidae), as subfamilies or (in the former case) even as tribe of subfamily Zopherinae. Some authors accept up to six subfamilies here, while others merge all except the Colydiinae into the Zopherinae.

The family has approximately 190 genera and 1700 species, which are found worldwide. A large number of members of the family feed on rotting wood or fungus associated with rotting wood, though some members of Colydiinae are predatory, or feed on living plant tissue such as roots, stems, flower stalks and fruit.^[1]

The oldest fossils of the family are Paleoendeitoma (subfamily Colydiinae, tribe Synchitini) and Cretomysteria from the early Late Cretaceous (Cenomanian) Burmese amber from Myanmar.^[2]

Systematics

Colydiinae (subfamily) Erichson, 1842^{i c g b}
- Acropini (tribe) Sharp, 1894^{c g}
  - Acropis (genus) Burmeister, 1840^{c g}
  - Plagiope Erichson, 1845^{c g}
- Adimerini Sharp, 1894^{c g}
  - Monoedus Horn, 1882^{i c g b}
- Colydiini Billberg, 1820^{c g b}
  - Anarmostes Pascoe, 1860^g
  - Aulonium Erichson, 1845^{i c g b}
  - Cicones Curtis, 1827^g
  - Colydium Fabricius, 1792^{i c g b}
  - Pseudaulonium Reitter, 1877^{c g}
- Gempylodini Sharp, 1893^{c g}
  - Gempylodes Pascoe, 1863^{c g}
- Nematidiini G.H. Horn, 1878^{c g}
  - Nematidium Erichson, 1845^{i c g b}
  - Phloeodalis Erichson, 1845^{c g}
- Ortherocerini Blanchard, 1845^{c g}
  - Orthocerus ^{c g}
- Rhagoderini G.H. Horn, 1878^{c g}
  - Rhagodera Mannerheim, 1843^{i c g b}
- Rhopalocerini Reitter, 1911^{c g}
  - Rhopalocerus W. Redtenbacher, 1842^{c g}
- Synchitini L. Redtenbacher, 1845^{c g}
  - Ablabus Broun, 1880^g
  - Bitoma Herbst, 1793^{i c g b}
  - Colobicus Latreille, 1807^{i c g b}
  - Corticus Germar, 1824^g
  - Coxelus Dejean, 1821^{i c g b}
  - Diodesma Latreille, 1829^g
  - Endophloeus Dejean, 1834^g
  - Langelandia Aubé, 1842^g
  - Lasconotus Erichson, 1845^{i c g b}
  - Lobogestoria Reitter, 1878^{i c g b}
  - Lyreus Aubé, 1861^{i c g b}
  - Megataphrus Casey, 1890^{i c g b}
  - Paha Dajoz, 1984^{i c g b}
  - Paxillobitoma Lord & Ivie, 2016^{c g}
  - Priolomus Erichson, 1845^{c g}
  - Pristoderus Hope, 1840^{c g}
  - Rechodes Erichson, 1845^{c g}
  - Semicoxelus Alekseev & Pankowski, 2020^{c g}
  - Synchita Hellwig, 1792^{c g}
  - Tarphiomimus Wollaston, 1862^{c g}
  - Tarphius Erichson, 1845^{c g}
  - Trachypholis Erichson, 1845^c
  - Xylolaemus Dejean, 1835^g
Zopherinae(subfamily) Solier, 1834^{i c g b}
- Latometini (tribe) Ślipiński & Lawrence, 1999^{g c}
  - Latometus (genus) Erichson, 1842^{c g}
- Monommatini Blanchard, 1845^{c g b}
  - Aspathines Champion, 1888^{i c g b}
  - Monomma Klug, 1833^{c g}
- Phellopsini LeConte, 1862^{i c g b}
  - Phellopsis LeConte, 1862^{i c g b}
- Pycnomerini Erichson, 1842^{c g b}
  - Pycnomerus Erichson, 1842^{i c g b}
- Usechini G. H. Horn, 1867^{c g b}
  - Usechus Motschulsky, 1845^{i c g b}
- Zopherini Solier, 1834^{c g b}
  - Zopherosis A. White, 1859^{c g}
  - Zopherus Laporte, 1840^{i c g b}
- Zopheromimini Alexeev & Nabhozhenko, 2023^{c g}
  - Zopheromimus Alexeev & Nabozhenko, 2023^{c g}

Data sources: i = ITIS,^[3] c = Catalogue of Life,^[4] g = GBIF,^[5] b = Bugguide.net^[6]

References

↑ Ślipinśki, Adam and Lawrence, John F.. "11.9. Zopheridae Solier, 1834". Volume 2 Morphology and Systematics (Elateroidea, Bostrichiformia, Cucujiformia partim), edited by Willy Kükenthal, Richard A.B. Leschen, Rolf G. Beutel and John F. Lawrence, Berlin, New York: De Gruyter, 2011, pp. 548-559
↑ Deng, Congshuang; Ślipiński, Adam; Ren, Dong; Pang, Hong (October 2017). "The first Mesozoic colydiid beetles (Coleoptera: Zopheridae: Colydiinae) from the Upper Cretaceous amber of Myanmar". Cretaceous Research. 78: 71–77. Bibcode:2017CrRes..78...71D. doi:10.1016/j.cretres.2017.04.010.
↑ "Zopheridae Report". Integrated Taxonomic Information System. Retrieved 2018-04-23.
↑ "Browse Zopheridae". Catalogue of Life. Archived from the original on 2021-10-03. Retrieved 2018-04-23.
↑ "Zopheridae". GBIF. Retrieved 2018-04-23.
↑ "Zopheridae Family Information". BugGuide.net. Retrieved 2018-04-23.

[1] Ślipinśki, Adam and Lawrence, John F.. "11.9. Zopheridae Solier, 1834". Volume 2 Morphology and Systematics (Elateroidea, Bostrichiformia, Cucujiformia partim), edited by Willy Kükenthal, Richard A.B. Leschen, Rolf G. Beutel and John F. Lawrence, Berlin, New York: De Gruyter, 2011, pp. 548-559

[2] Deng, Congshuang; Ślipiński, Adam; Ren, Dong; Pang, Hong (October 2017). "The first Mesozoic colydiid beetles (Coleoptera: Zopheridae: Colydiinae) from the Upper Cretaceous amber of Myanmar". Cretaceous Research. 78: 71–77. Bibcode:2017CrRes..78...71D. doi:10.1016/j.cretres.2017.04.010.

[itis-3] "Zopheridae Report". Integrated Taxonomic Information System. Retrieved 2018-04-23.

[catlife-4] "Browse Zopheridae". Catalogue of Life. Archived from the original on 2021-10-03. Retrieved 2018-04-23.

[gbif-5] "Zopheridae". GBIF. Retrieved 2018-04-23.

[bugref-6] "Zopheridae Family Information". BugGuide.net. Retrieved 2018-04-23.

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// Workers AI · dad joke modeWhat did the Zopheridae beetle say? "I'm beetle down with work.

See also

Systematics

References