// Workers AI · dad joke modeWhat did Coryphella say? Sea you later.
| Coryphella | |
|---|---|
| Coryphella verrucosa | |
| Scientific classification | |
| Kingdom: | Animalia |
| Phylum: | Mollusca |
| Class: | Gastropoda |
| Order: | Nudibranchia |
| Suborder: | Aeolidacea |
| Superfamily: | Flabellinoidea |
| Family: | Flabellinidae Bergh, 1889[1] |
| Genus: | Coryphella M.E. Gray, 1850[2] |
| Synonyms | |
|
List
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Coryphella is a genus of sea slugs, specifically aeolid nudibranchs, marine gastropod molluscs in the family Flabellinidae.[3][4]
Taxonomy
[edit]Coryphella was originally described in an 1850 paper, and included three original species; Coryphella landsburgii, Coryphella rufobranchialis, and Coryphella pellucida.[2] The family Coryphellidae was first established in 1889 by Rudolph Bergh,[1] and was re-established in a 2017 study.[5] Molecular studies have found that many existing genera were actually synonymous with Coryphella, leading to many genera being merged.[6] However, these studies had failed to take morphological analysis into account, the "lumping" of several genera under Coryphella leading to loss of differentiation. Thus in Korshunova et al. (2025)[7], after full molecular and morphological analysis, the genus Coryphella was restricted, previously synonymized genera reinstated, and two new genera established. The most recent rebuttal to Korshunova et al., published by Ekimova et al. (2026), re-lumped the reinstated genera, along with the newly established genera, back to Coryphella[3].
Species
[edit]According to Korshunova et al. (2025), species within the genus Coryphella are as follows[7]:
- Coryphella longicaudata O'Donoghue, 1922
- Coryphella pseudoverrucosa Martynov, Sanamyan & Korshunova, 2015
- Coryphella verrucosa M. Sars, 1829
Genus incertae sedis under Coryphella according to Korshunova et al. (2025) include[7]:
- Coryphella abei Baba, 1987
- Coryphella alderi Adams, 1861
- Coryphella pallida A. E. Verrill, 1900
- Coryphella subrosacea Eschscholtz, 1831
According to Ekimova et al. (2026), species with the genus Coryphella are as follows[3]:
- Coryphella abei Baba, 1987
- Coryphella alexanderi Ekimova, 2022
- Coryphella amabilis (Hirano & Kuzirian, 1991)
- Coryphella athadona Bergh, 1875
- Coryphella borealis Odhner, 1922
- Coryphella browni Picton, 1980
- Coryphella californica Bergh, 1904
- Coryphella candela (Korshunova, Fletcher & Martynov, 2025)
- Coryphella capensis Thiele, 1925
- Coryphella cerverai (M. A. Fischer, van der Velde & Roubos, 2007)
- Coryphella chriskaugei (Korshunova, Martynov, Bakken, Evertsen, Fletcher, Mudianta, Saito, Lundin, Schrödl & Picton, 2017)
- Coryphella cooperi Cockerell, 1901
- Coryphella dana (Millen & Hamann, 2006)
- Coryphella falklandica Eliot, 1907
- Coryphella fogata (Millen & Hermosillo, 2007)
- Coryphella gracilis (Alder & Hancock, 1844)
- Coryphella insolita (García-Gómez & Cervera, 1989)
- Coryphella lineata (Lovén, 1846)
- Coryphella longicaudata O'Donoghue, 1922
- Coryphella monicae (Korshunova, Martynov, Bakken, Evertsen, Fletcher, Mudianta, Saito, Lundin, Schrödl & Picton, 2017)
- Coryphella nobilis A. E. Verrill, 1880
- Coryphella orjani (Korshunova, Martynov, Bakken, Evertsen, Fletcher, Mudianta, Saito, Lundin, Schrödl & Picton, 2017)
- Coryphella pallida A. E. Verrill, 1900
- Coryphella pseudoverrucosa Martynov, Sanamyan & Korshunova, 2015
- Coryphella sanamyanae (Korshunova, Martynov, Bakken, Evertsen, Fletcher, Mudianta, Saito, Lundin, Schrödl & Picton, 2017)
- Coryphella trilineata O'Donoghue, 1921
- Coryphella trophina (Bergh, 1890)'
- Coryphella verrucosa (M. Sars, 1829)
- Coryphella verta Ev. Marcus, 1970
It is very important to note that despite Orienthella being invalidated, Orienthella piunca currently remains an accepted species, as it was the only member of Orienthella not studied in Ekimova et al. 2026.
- Coryphella verrucosa, spawn, Gulen Dive Resort, Norway
- Coryphella verrucosa, Gulen Dive Resort, Norway
- Coryphella longicaudata, British Columbia
- Coryphella longicaudata, Kenai Fjords national Park, Alaska
Species brought into synonymy following Ekimova et al. (2026):[3][4]:
- Coryphella albomarginata M. C. Miller, 1971: synonym of Coryphellina albomarginata (M. C. Miller, 1971)
- Coryphella beaumonti Eliot, 1906: synonym of Cumanotus beaumonti (Eliot, 1906)
- Coryphella bostoniensis (Couthouy, 1838): synonym of Facelina bostoniensis (Couthouy, 1838)
- Coryphella cynara Ev. Marcus & Er. Marcus, 1967: synonym of Kynaria cynara (Ev. Marcus & Er. Marcus, 1967)
- Coryphella dushia Ev. Marcus & Er. Marcus, 1963: synonym of Edmundsella dushia (Ev. Marcus & Er. Marcus, 1963)
- Coryphella fisheri MacFarland, 1966: synonym of accepted as Orienthella piunca (Er. Marcus, 1961)
- Coryphella fusca O'Donoghue, 1921: synonym of Coryphella trophina (Bergh, 1890)
- Coryphella islandica Odhner, 1937: synonym of Chlamylla islandica (Odhner, 1937)
- Coryphella japonica Volodchenko, 1941: synonym of Ziminella japonica Volodchenko, 1941
- Coryphella laevidens Knipowitsch, 1902: synonym of Ziminella salmonacea (Couthouy, 1838)
- Coryphella mananensis (W. Stimpson, 1853): synonym of Coryphella verrucosa (M. Sars, 1829)
- Coryphella ornata Risbec, 1928: synonym of Samla bicolor (Kelaart, 1858)
- Coryphella parva M. Hadfield, 1963: synonym of Chlamylla parva {M. Hadfield, 1963
- Coryphella pedata (Montagu, 1816): synonym of Edmundsella pedata (Montagu, 1816)
- Coryphella pellucida (Alder & Hancock, 1843): synonym of Flabellina pellucida (Alder & Hancock, 1843)
- Coryphella piunca Er. Marcus, 1981: synonym of Orienthella piunca (Er. Marcus, 1961)
- Coryphella polaris Volodchenko, 1946: synonym of Chlamylla polaris (Volodchenko, 1946)
- Coryphella pricei MacFarland, 1966: synonym of Apata pricei (MacFarland, 1966)
- Coryphella robusta Trinchese, 1874: synonym of Coryphella verrucosa (M. Sars, 1829)
- Coryphella rufibranchialis (G. Johnston, 1832): synonym of Coryphella verrucosa (M. Sars, 1829)
- Coryphella rutila Verrill, 1879: synonym of Flabellina pellucida (Alder & Hancock, 1843)
- Coryphella salmonacea (Couthouy, 1838): synonym of Ziminella salmonacea (Couthouy, 1838)
- Coryphella sarsi Friele, 1903: synonym of Coryphella nobilis A. E. Verrill, 1880
- Coryphella stimpsoni (A. E. Verrill, 1879): synonym of Ziminella salmonacea (Couthouy, 1838)
References
[edit]- 1 2 Winckworth, R. (21 May 1946). "On Bergh's Malacologische Untersuchungen". Proceedings of the Malacological Society of London. 27 (1): 1038 – via Biodiversity Heritage Library.
- 1 2 Gray, Maria Emma; Gray, John Edward (1859). Figures of molluscous animals, selected from various authors. Vol. 4. London: Longman, Brown, Green and Longmans. p. 109. doi:10.5962/bhl.title.10461. OCLC 9597391 – via Biodiversity Heritage Library.
- 1 2 3 4 Ekimova I., Carmona L., Mikhlina AL., Grishina D., Stanovova M.V., Schepetov D.M., Hoover C., de Souza-Canal J., Kuznetsov K.O., Valdés A.; (2026). Neither "lumpers" nor "splitters": A global revision of Flabellinidae s.l. nudibranchs (Gastropoda: Heterobranchia: Nudibranchia). PLoS One 21(5): e0347759.
- 1 2 Bouchet, P. (2019) Coryphella J. E. Gray, 1850. MolluscaBase eds. (2026). MolluscaBase. Accessed through: World Register of Marine Species on 2026-05-05
- ↑ Korshunova, T.; Martynov, A.; Bakken, T.; Evertsen, J.; Fletcher, K.; Mudianta, W.; Saito, H.; Lundin, K.; Schrödl, M.; Picton, B. (2017). Polyphyly of the traditional family Flabellinidae affects a major group of Nudibranchia: aeolidacean taxonomic reassessment with descriptions of several new families, genera, and species (Mollusca, Gastropoda). ZooKeys. 717: 1-139.
- ↑ Ekimova, Irina; Valdés, Ángel; Malaquias, Manuel António E; Rauch, Cessa; Chichvarkhin, Anton; Mikhlina, Anna; Antokhina, Tatiana; Chichvarkhina, Olga; Schepetov, Dimitry (16 March 2022). "High-level taxonomic splitting in allopatric taxa causes confusion downstream: a revision of the nudibranch family Сoryphellidae". Zoological Journal of the Linnean Society. 196 (1): 215–249. doi:10.1093/zoolinnean/zlab109.
- 1 2 3 Korshunova T.; Fletcher K.; Martynov A. (2025). The endless forms are the most differentiated—how taxonomic pseudo-optimization masked natural diversity and evolution: the nudibranch case Zoological Journal of the Linnean Society. Volume 204, Issue 4, August 2025.